HoldingsChannel.com
Funds Holding IONS
HolderAmountPosition Size
($ in 1000's)
 As of
FMR LLC20,960,227$990,999 03/31/2020
     IONS20,960,227$990,999 03/31/2020
Vanguard Group Inc.12,053,561$569,893 03/31/2020
     IONS12,053,561$569,893 03/31/2020
Baillie Gifford & Co.8,745,683$413,496 03/31/2020
     IONS8,745,683$413,496 03/31/2020
Bellevue Group AG8,379,642$396,189 03/31/2020
     IONS8,379,642$396,189 03/31/2020
BlackRock Inc.8,167,940$386,180 03/31/2020
     IONS8,167,940$386,180 03/31/2020
Clearbridge Investments LLC8,034,264$379,860 03/31/2020
     IONS8,034,264$379,860 03/31/2020
State Street Corp $200,082 03/31/2020
     IONS3,892,894$184,056 03/31/2020
     NOTE 115,740,000$16,026 03/31/2020
Wellington Management Group LLP3,985,217$188,422 03/31/2020
     IONS3,985,217$188,422 03/31/2020
Price T Rowe Associates Inc. MD3,762,445$177,888 03/31/2020
     IONS3,762,445$177,888 03/31/2020
Invesco Ltd.2,340,797$110,673 03/31/2020
     IONS2,340,797$110,673 03/31/2020
Waddell & Reed Financial Inc.2,166,311$102,423 03/31/2020
     IONS2,166,311$102,423 03/31/2020
BVF Inc. IL1,944,517$91,937 03/31/2020
     IONS1,944,517$91,937 03/31/2020
Citadel Advisors LLC $83,420 03/31/2020
     IONS1,738,992$82,219 03/31/2020
     Call65,700$3,106 03/31/2020
     Put40,300$1,905 03/31/2020
Renaissance Technologies LLC1,688,411$79,828 03/31/2020
     IONS1,688,411$79,828 03/31/2020
Geode Capital Management LLC1,491,065$70,439 03/31/2020
     IONS1,491,065$70,439 03/31/2020
American Century Companies Inc.1,407,628$66,553 03/31/2020
     IONS1,407,628$66,553 03/31/2020
Two Sigma Advisers LP1,187,211$56,131 03/31/2020
     IONS1,187,211$56,131 03/31/2020
Wells Fargo & Company MN967,293$45,733 03/31/2020
     IONS967,293$45,733 03/31/2020
First Trust Advisors LP952,473$45,033 03/31/2020
     IONS952,473$45,033 03/31/2020
Sumitomo Mitsui Trust Holdings Inc.921,187$43,554 03/31/2020
     IONS921,187$43,554 03/31/2020
Northern Trust Corp835,493$39,501 03/31/2020
     IONS835,493$39,501 03/31/2020
Bank of America Corp DE745,575$35,251 03/31/2020
     IONS745,575$35,251 03/31/2020
Goldman Sachs Group Inc. $34,030 03/31/2020
     IONS719,750$34,030 03/31/2020
     Put20,000$946 03/31/2020
     Call20,000$946 03/31/2020
Dowling & Yahnke LLC675,478$31,937 03/31/2020
     IONS675,478$31,937 03/31/2020
Rockefeller Capital Management L.P.670,138$31,684 03/31/2020
     IONS670,138$31,684 03/31/2020
Pinnacle Associates Ltd.621,909$29,404 03/31/2020
     IONS621,909$29,404 03/31/2020
Charles Schwab Investment Management Inc.610,699$28,874 03/31/2020
     IONS610,699$28,874 03/31/2020
Bank of New York Mellon Corp553,970$26,191 03/31/2020
     IONS553,970$26,191 03/31/2020
Two Sigma Investments LP547,997$25,909 03/31/2020
     IONS547,997$25,909 03/31/2020
Swiss National Bank518,759$24,527 03/31/2020
     IONS518,759$24,527 03/31/2020
Nuveen Asset Management LLC477,227$22,564 03/31/2020
     IONS477,227$22,564 03/31/2020
Mitsubishi UFJ Trust & Banking Corp467,730$22,114 03/31/2020
     IONS467,730$22,114 03/31/2020
Legal & General Group Plc435,038$20,629 03/31/2020
     IONS435,038$20,629 03/31/2020
Sicart Associates LLC429,752$20,319 03/31/2020
     IONS429,752$20,319 03/31/2020
Stifel Financial Corp390,696$18,488 03/31/2020
     IONS390,696$18,488 03/31/2020
Susquehanna International Group LLP $18,302 03/31/2020
     IONS378,493$17,895 03/31/2020
     Call279,400$13,210 03/31/2020
     Put270,800$12,803 03/31/2020
Nikko Asset Management Americas Inc.381,112$18,019 03/31/2020
     IONS381,112$18,019 03/31/2020
UBS Asset Management Americas Inc.360,732$17,055 03/31/2020
     IONS360,732$17,055 03/31/2020
Raymond James & Associates354,504$16,761 03/31/2020
     IONS354,504$16,761 03/31/2020
Bank of Montreal Can340,444$16,684 03/31/2020
     IONS340,444$16,684 03/31/2020
Altrinsic Global Advisors LLC348,834$16,493 03/31/2020
     IONS348,834$16,493 03/31/2020
Dimensional Fund Advisors LP348,572$16,482 03/31/2020
     IONS348,572$16,482 03/31/2020
Rothschild & Co. Asset Management US Inc.344,027$16,266 03/31/2020
     IONS344,027$16,266 03/31/2020
Candriam Luxembourg S.C.A.261,700$15,454 03/31/2020
     IONS261,700$15,454 03/31/2020
California Public Employees Retirement System286,553$13,548 03/31/2020
     IONS286,553$13,548 03/31/2020
Tocqueville Asset Management L.P.284,642$13,458 03/31/2020
     IONS284,642$13,458 03/31/2020
Sector Gamma AS275,481$13,025 03/31/2020
     IONS275,481$13,025 03/31/2020
Bank Julius Baer & Co. Ltd Zurich274,081$12,945 03/31/2020
     IONS274,081$12,945 03/31/2020
JPMorgan Chase & Co.271,722$12,847 03/31/2020
     IONS271,722$12,847 03/31/2020
New York State Common Retirement Fund271,531$12,838 03/31/2020
     IONS271,531$12,838 03/31/2020
Franklin Resources Inc.240,673$11,379 03/31/2020
     IONS240,673$11,379 03/31/2020
Peregrine Capital Management LLC222,342$10,512 03/31/2020
     IONS222,342$10,512 03/31/2020
California State Teachers Retirement System222,147$10,503 03/31/2020
     IONS222,147$10,503 03/31/2020
EcoR1 Capital LLC208,609$9,863 03/31/2020
     IONS208,609$9,863 03/31/2020
Parametric Portfolio Associates LLC204,824$9,684 03/31/2020
     IONS204,824$9,684 03/31/2020
Morgan Stanley202,748$9,587 03/31/2020
     IONS202,748$9,587 03/31/2020
First Light Asset Management LLC195,850$9,260 03/31/2020
     IONS195,850$9,260 03/31/2020
Tekla Capital Management LLC193,201$9,135 03/31/2020
     IONS193,201$9,135 03/31/2020
APG Asset Management N.V.183,160$8,660 03/31/2020
     IONS183,160$8,660 03/31/2020
BLB&B Advisors LLC180,245$8,522 03/31/2020
     IONS180,245$8,522 03/31/2020
State Board of Administration of Florida Retirement System176,004$8,321 03/31/2020
     IONS176,004$8,321 03/31/2020
Citigroup Inc. $8,025 03/31/2020
     IONS175,338$8,290 03/31/2020
     Put14,800$700 03/31/2020
     Call9,200$435 03/31/2020
Royal Bank of Canada160,652$7,595 03/31/2020
     IONS160,652$7,595 03/31/2020
Squarepoint Ops LLC148,144$7,004 03/31/2020
     IONS148,144$7,004 03/31/2020
Deutsche Bank AG148,130$7,002 03/31/2020
     IONS148,130$7,002 03/31/2020
Natixis Advisors L.P.139,707$6,605 03/31/2020
     IONS139,707$6,605 03/31/2020
Rhumbline Advisers139,451$6,593 03/31/2020
     IONS139,451$6,593 03/31/2020
DAVENPORT & Co LLC136,826$6,469 03/31/2020
     IONS136,826$6,469 03/31/2020
Credit Suisse AG134,631$6,365 03/31/2020
     IONS134,631$6,365 03/31/2020
ARK Investment Management LLC119,681$5,659 03/31/2020
     IONS119,681$5,659 03/31/2020
Edmond DE Rothschild Holding S.A.113,750$5,378 03/31/2020
     IONS113,750$5,378 03/31/2020
Sonora Investment Management LLC112,262$5,308 03/31/2020
     IONS112,262$5,308 03/31/2020
Advent Capital Management DE $5,301 03/31/2020
     CONV5,283,000$5,301 03/31/2020
Lazard Asset Management LLC $5,196 03/31/2020
     Bond 30 360P2 U5,000,000$5,072 03/31/2020
     IONS2,639$124 03/31/2020
Jacobs Levy Equity Management Inc.105,577$4,992 03/31/2020
     IONS105,577$4,992 03/31/2020
State of New Jersey Common Pension Fund D101,352$4,792 03/31/2020
     IONS101,352$4,792 03/31/2020
Gotham Asset Management LLC97,992$4,633 03/31/2020
     IONS97,992$4,633 03/31/2020
Andra AP fonden98,000$4,633 03/31/2020
     IONS98,000$4,633 03/31/2020
TD Asset Management Inc.95,765$4,528 03/31/2020
     IONS95,765$4,528 03/31/2020
Canada Pension Plan Investment Board92,951$4,395 03/31/2020
     IONS92,951$4,395 03/31/2020
UBS Group AG91,103$4,308 03/31/2020
     IONS91,103$4,308 03/31/2020
Sei Investments Co.90,483$4,278 03/31/2020
     IONS90,483$4,278 03/31/2020
AXA88,844$4,201 03/31/2020
     IONS88,844$4,201 03/31/2020
Barclays PLC $4,189 03/31/2020
     IONS62,822$2,970 03/31/2020
     Call52,800$2,496 03/31/2020
     Put27,000$1,277 03/31/2020
Skandinaviska Enskilda Banken AB publ 87,400$4,132 03/31/2020
     IONS87,400$4,132 03/31/2020
Majedie Asset Management Ltd82,377$3,904 03/31/2020
     IONS82,377$3,904 03/31/2020
WINTON GROUP Ltd82,360$3,894 03/31/2020
     IONS82,360$3,894 03/31/2020
Van ECK Associates Corp81,766$3,866 03/31/2020
     IONS81,766$3,866 03/31/2020
Gateway Investment Advisers LLC76,476$3,616 03/31/2020
     IONS76,476$3,616 03/31/2020
Los Angeles Capital Management & Equity Research Inc.76,228$3,604 03/31/2020
     IONS76,228$3,604 03/31/2020
Aviva PLC75,733$3,581 03/31/2020
     IONS75,733$3,581 03/31/2020
Capital Impact Advisors LLC72,267$3,417 03/31/2020
     IONS72,267$3,417 03/31/2020
Millennium Management LLC71,592$3,385 03/31/2020
     IONS71,592$3,385 03/31/2020
State of Wisconsin Investment Board70,176$3,318 03/31/2020
     IONS70,176$3,318 03/31/2020
Amia Capital LLP69,678$3,294 03/31/2020
     IONS69,678$3,294 03/31/2020
Alliancebernstein L.P.68,958$3,260 03/31/2020
     IONS68,958$3,260 03/31/2020
Guggenheim Capital LLC65,890$3,115 03/31/2020
     IONS65,890$3,115 03/31/2020
Asset Management One Co. Ltd.64,607$3,054 03/31/2020
     IONS64,607$3,054 03/31/2020
D. E. Shaw & Co. Inc. $3,021 03/31/2020
     Call38,900$1,839 03/31/2020
     IONS24,996$1,182 03/31/2020
Russell Investments Group Ltd.62,087$2,939 03/31/2020
     IONS62,087$2,939 03/31/2020
Treasurer of the State of North Carolina60,980$2,883 03/31/2020
     IONS60,980$2,883 03/31/2020
Federated Hermes Inc.58,235$2,753 03/31/2020
     IONS58,235$2,753 03/31/2020
Envestnet Asset Management Inc.55,610$2,629 03/31/2020
     IONS55,610$2,629 03/31/2020
International Biotechnology Trust PLC53,781$2,543 03/31/2020
     IONS53,781$2,543 03/31/2020
Public Employees Retirement System of Ohio51,572$2,438 03/31/2020
     IONS51,572$2,438 03/31/2020
Pictet Asset Management Ltd.49,700$2,350 03/31/2020
     IONS49,700$2,350 03/31/2020
Fjarde AP Fonden Fourth Swedish National Pension Fund49,300$2,331 03/31/2020
     IONS49,300$2,331 03/31/2020
Trexquant Investment LP47,499$2,246 03/31/2020
     IONS47,499$2,246 03/31/2020
Janus Henderson Group PLC47,274$2,236 03/31/2020
     IONS47,274$2,236 03/31/2020
LPL Financial LLC46,848$2,215 03/31/2020
     IONS46,848$2,215 03/31/2020
Voya Investment Management LLC45,756$2,163 03/31/2020
     IONS45,756$2,163 03/31/2020
Hudson Bay Capital Management LP45,000$2,128 03/31/2020
     IONS45,000$2,128 03/31/2020
Optimal Asset Management Inc.43,575$2,060 03/31/2020
     IONS43,575$2,060 03/31/2020
Janney Montgomery Scott LLC39,980$1,890 03/31/2020
     IONS39,980$1,890 03/31/2020
Capital Counsel LLC NY39,380$1,862 03/31/2020
     IONS39,380$1,862 03/31/2020
Cubist Systematic Strategies LLC37,907$1,792 03/31/2020
     IONS37,907$1,792 03/31/2020
Principal Financial Group Inc.36,702$1,735 03/31/2020
     IONS36,702$1,735 03/31/2020
Bridgeway Capital Management Inc.36,200$1,712 03/31/2020
     IONS36,200$1,712 03/31/2020
PFS Investments Inc.36,163$1,710 03/31/2020
     IONS36,163$1,710 03/31/2020
L2 Asset Management LLC36,129$1,708 03/31/2020
     IONS36,129$1,708 03/31/2020
Aigen Investment Management LP35,254$1,667 03/31/2020
     IONS35,254$1,667 03/31/2020
Amalgamated Bank33,705$1,594 03/31/2020
     IONS33,705$1,594 03/31/2020
Great West Life Assurance Co. Can33,144$1,578 03/31/2020
     IONS33,144$1,578 03/31/2020
Korea Investment CORP33,200$1,570 03/31/2020
     IONS33,200$1,570 03/31/2020
Capital Fund Management S.A.33,111$1,565 03/31/2020
     IONS33,111$1,565 03/31/2020
Employees Retirement System of Texas32,409$1,532 03/31/2020
     IONS32,409$1,532 03/31/2020
AJO LP31,831$1,504 03/31/2020
     IONS31,831$1,504 03/31/2020
AMP Capital Investors Ltd30,948$1,491 03/31/2020
     IONS30,948$1,491 03/31/2020
AustralianSuper Pty Ltd31,060$1,469 03/31/2020
     IONS31,060$1,469 03/31/2020
Arizona State Retirement System30,752$1,454 03/31/2020
     IONS30,752$1,454 03/31/2020
Macquarie Group Ltd.30,602$1,447 03/31/2020
     IONS30,602$1,447 03/31/2020
Group One Trading L.P. $1,357 03/31/2020
     Call48,300$2,284 03/31/2020
     Put19,600$927 03/31/2020
Circle Wealth Management LLC27,900$1,319 03/31/2020
     IONS27,900$1,319 03/31/2020
Aperio Group LLC27,530$1,301 03/31/2020
     IONS27,530$1,301 03/31/2020
Amundi Pioneer Asset Management Inc.27,065$1,281 03/31/2020
     IONS27,065$1,281 03/31/2020
Hussman Strategic Advisors Inc.25,000$1,182 03/31/2020
     IONS25,000$1,182 03/31/2020
Utah Retirement Systems24,409$1,154 03/31/2020
     IONS24,409$1,154 03/31/2020
Profund Advisors LLC24,051$1,137 03/31/2020
     IONS24,051$1,137 03/31/2020
Zurcher Kantonalbank Zurich Cantonalbank 22,730$1,075 03/31/2020
     IONS22,730$1,075 03/31/2020
Nomura Asset Management Co. Ltd.22,600$1,069 03/31/2020
     IONS22,600$1,069 03/31/2020
Stephens Inc. AR22,311$1,055 03/31/2020
     IONS22,311$1,055 03/31/2020
Xact Kapitalforvaltning AB22,311$1,055 03/31/2020
     IONS22,311$1,055 03/31/2020
Mitsubishi UFJ Kokusai Asset Management Co. Ltd.22,166$1,048 03/31/2020
     IONS22,166$1,048 03/31/2020
Vestmark Advisory Solutions Inc.21,827$1,032 03/31/2020
     IONS21,827$1,032 03/31/2020
ProShare Advisors LLC21,707$1,026 03/31/2020
     IONS21,707$1,026 03/31/2020
Oaktree Capital Management LP $1,014 03/31/2020
     CONVERTIBLE BOND1,000,000$1,014 03/31/2020
AlphaCrest Capital Management LLC20,881$987 03/31/2020
     IONS20,881$987 03/31/2020
Tanaka Capital Management Inc.20,040$947 03/31/2020
     IONS20,040$947 03/31/2020
New England Research & Management Inc.20,040$947 03/31/2020
     IONS20,040$947 03/31/2020
Comerica Bank16,279$946 03/31/2020
     IONS16,279$946 03/31/2020
BOKF NA19,718$932 03/31/2020
     IONS19,718$932 03/31/2020
Eaton Vance Management19,659$929 03/31/2020
     IONS19,659$929 03/31/2020
Public Employees Retirement Association of Colorado19,604$927 03/31/2020
     IONS19,604$927 03/31/2020
Commonwealth Equity Services LLC19,521$922 03/31/2020
     IONS19,521$922 03/31/2020
Creative Planning19,246$910 03/31/2020
     IONS19,246$910 03/31/2020
MERIAN GLOBAL INVESTORS UK Ltd18,484$873 03/31/2020
     IONS18,484$873 03/31/2020
Reynders McVeigh Capital Management LLC18,390$869 03/31/2020
     IONS18,390$869 03/31/2020
Arden Trust Co18,345$867 03/31/2020
     IONS18,345$867 03/31/2020
Toronto Dominion Bank18,235$863 03/31/2020
     IONS18,235$863 03/31/2020
National Asset Management Inc.18,218$861 03/31/2020
     IONS18,218$861 03/31/2020
Alken Asset Management Ltd. $848 03/31/2020
     US DOMESTIC850,000$848 03/31/2020
Alexandria Capital LLC17,908$847 03/31/2020
     IONS17,908$847 03/31/2020
Advisors Asset Management Inc.17,833$843 03/31/2020
     IONS17,833$843 03/31/2020
Victory Capital Management Inc.17,737$839 03/31/2020
     IONS17,737$839 03/31/2020
Lincoln National Corp17,572$831 03/31/2020
     IONS17,572$831 03/31/2020
Grandfield & Dodd LLC17,336$820 03/31/2020
     IONS17,336$820 03/31/2020
Starfire Investment Advisers Inc.16,543$782 03/31/2020
     IONS16,543$782 03/31/2020
Castle Rock Wealth Management LLC14,863$778 03/31/2020
     IONS14,863$778 03/31/2020
Jane Street Group LLC $765 03/31/2020
     Call38,100$1,801 03/31/2020
     Put30,400$1,437 03/31/2020
     IONS8,476$401 03/31/2020
Blair William & Co. IL16,111$762 03/31/2020
     IONS16,111$762 03/31/2020
Raymond James Financial Services Advisors Inc.16,089$761 03/31/2020
     IONS16,089$761 03/31/2020
Teacher Retirement System of Texas15,529$734 03/31/2020
     IONS15,529$734 03/31/2020
Chicago Equity Partners LLC15,140$716 03/31/2020
     IONS15,140$716 03/31/2020
Cowen AND Company LLC $704 03/31/2020
     IONS20,797$983 03/31/2020
     Put27,700$292 03/31/2020
     Call2,000$13 03/31/2020
AGF Investments LLC14,827$701 03/31/2020
     IONS14,827$701 03/31/2020
BRITISH COLUMBIA INVESTMENT MANAGEMENT Corp14,618$691 03/31/2020
     IONS14,618$691 03/31/2020
Shell Asset Management Co.13,795$652 03/31/2020
     IONS13,795$652 03/31/2020
North Star Asset Management Inc.13,700$648 03/31/2020
     IONS13,700$648 03/31/2020
Advisor Group Holdings Inc.13,577$643 03/31/2020
     IONS13,577$643 03/31/2020
DNB Asset Management AS13,226$625 03/31/2020
     IONS13,226$625 03/31/2020
M&T Bank Corp12,759$603 03/31/2020
     IONS12,759$603 03/31/2020
Wrapmanager Inc.12,582$595 03/31/2020
     IONS12,582$595 03/31/2020
SG Americas Securities LLC12,503$591 03/31/2020
     IONS12,503$591 03/31/2020
Schonfeld Strategic Advisors LLC12,436$588 03/31/2020
     IONS12,436$588 03/31/2020
Oppenheimer Asset Management Inc.12,223$577 03/31/2020
     IONS12,223$577 03/31/2020
HighTower Advisors LLC12,178$570 03/31/2020
     IONS12,178$570 03/31/2020
Nissay Asset Management Corp Japan ADV12,010$568 03/31/2020
     IONS12,010$568 03/31/2020
Horizon Kinetics Asset Management LLC12,000$567 03/31/2020
     IONS12,000$567 03/31/2020
NJ State Employees Deferred Compensation Plan12,000$567 03/31/2020
     IONS12,000$567 03/31/2020
IFM Investors Pty Ltd11,692$564 03/31/2020
     IONS11,692$564 03/31/2020
Truist Financial Corp11,672$551 03/31/2020
     IONS11,672$551 03/31/2020
Tudor Investment Corp Et Al11,546$546 03/31/2020
     IONS11,546$546 03/31/2020
Berkshire Asset Management LLC PA11,516$544 03/31/2020
     IONS11,516$544 03/31/2020
Hsbc Holdings PLC11,314$536 03/31/2020
     IONS11,314$536 03/31/2020
Piedmont Investment Advisors Inc.11,233$531 03/31/2020
     IONS11,233$531 03/31/2020
Qube Research & Technologies Ltd11,225$531 03/31/2020
     IONS11,225$531 03/31/2020
Fifth Third Bancorp11,188$529 03/31/2020
     IONS11,188$529 03/31/2020
KBC Group NV10,971$519 03/31/2020
     IONS10,971$519 03/31/2020
361 Capital LLC10,823$512 03/31/2020
     IONS10,823$512 03/31/2020
Seeyond4,429$509 03/31/2020
     IONS4,429$509 03/31/2020
Wolverine Trading LLC $502 03/31/2020
     Put113,400$5,467 03/31/2020
     IONS100,517$4,846 03/31/2020
     Call23,300$1,123 03/31/2020
Westpac Banking Corp10,594$501 03/31/2020
     IONS10,594$501 03/31/2020
Coastal Investment Advisors Inc.10,037$475 03/31/2020
     IONS10,037$475 03/31/2020
Nicollet Investment Management Inc.10,005$473 03/31/2020
     IONS10,005$473 03/31/2020
Ameriprise Financial Inc.9,960$472 03/31/2020
     IONS9,960$472 03/31/2020
Nisa Investment Advisors LLC9,721$460 03/31/2020
     IONS9,721$460 03/31/2020
Man Group plc9,700$459 03/31/2020
     IONS9,700$459 03/31/2020
Virtus ETF Advisers LLC9,329$441 03/31/2020
     IONS9,329$441 03/31/2020
Metropolitan Life Insurance Co NY8,973$424 03/31/2020
     IONS8,973$424 03/31/2020
PNC Financial Services Group Inc.8,584$405 03/31/2020
     IONS8,584$405 03/31/2020
DekaBank Deutsche Girozentrale7,899$376 03/31/2020
     IONS7,899$376 03/31/2020
First Quadrant L P CA7,939$375 03/31/2020
     IONS7,939$375 03/31/2020
Standard Life Aberdeen plc7,851$372 03/31/2020
     IONS7,851$372 03/31/2020
Prospera Financial Services Inc7,850$371 03/31/2020
     IONS7,850$371 03/31/2020
Northeast Financial Consultants Inc7,740$366 03/31/2020
     IONS7,740$366 03/31/2020
Keybank National Association OH7,617$360 03/31/2020
     IONS7,617$360 03/31/2020
Engineers Gate Manager LP7,598$359 03/31/2020
     IONS7,598$359 03/31/2020
Prudential Financial Inc.7,501$355 03/31/2020
     IONS7,501$355 03/31/2020
Rafferty Asset Management LLC7,494$354 03/31/2020
     IONS7,494$354 03/31/2020
Tower Research Capital LLC TRC 7,482$354 03/31/2020
     IONS7,482$354 03/31/2020
Sumitomo Mitsui DS Asset Management Company Ltd7,236$342 03/31/2020
     IONS7,236$342 03/31/2020
Caisse DE Depot ET Placement DU Quebec7,166$339 03/31/2020
     IONS7,166$339 03/31/2020
Teza Capital Management LLC7,168$339 03/31/2020
     IONS7,168$339 03/31/2020
Rathbone Brothers plc6,945$328 03/31/2020
     IONS6,945$328 03/31/2020
Commerce Bank6,693$316 03/31/2020
     IONS6,693$316 03/31/2020
Daiwa Securities Group Inc.6,248$295 03/31/2020
     IONS6,248$295 03/31/2020
ETRADE Capital Management LLC6,121$289 03/31/2020
     IONS6,121$289 03/31/2020
ExodusPoint Capital Management LP5,934$281 03/31/2020
     IONS5,934$281 03/31/2020
AQR Capital Management LLC5,905$279 03/31/2020
     IONS5,905$279 03/31/2020
FDx Advisors Inc.5,727$271 03/31/2020
     IONS5,727$271 03/31/2020
First Republic Investment Management Inc.5,616$266 03/31/2020
     IONS5,616$266 03/31/2020
Point72 Asset Management L.P.5,573$263 03/31/2020
     IONS5,573$263 03/31/2020
Baird Financial Group Inc.5,559$263 03/31/2020
     IONS5,559$263 03/31/2020
AGF Investments Inc.5,502$260 03/31/2020
     IONS5,502$260 03/31/2020
Bayesian Capital Management LP5,300$251 03/31/2020
     IONS5,300$251 03/31/2020
Nordea Investment Management AB5,156$247 03/31/2020
     IONS5,156$247 03/31/2020
Brown Advisory Securities LLC4,800$247 03/31/2020
     IONS4,800$247 03/31/2020
Buckingham Asset Management LLC5,212$246 03/31/2020
     IONS5,212$246 03/31/2020
Exchange Traded Concepts LLC5,184$245 03/31/2020
     IONS5,184$245 03/31/2020
Advisory Services Network LLC5,183$245 03/31/2020
     IONS5,183$245 03/31/2020
CIBC Asset Management Inc5,116$242 03/31/2020
     IONS5,116$242 03/31/2020
Robeco Institutional Asset Management B.V.5,038$238 03/31/2020
     IONS5,038$238 03/31/2020
Whitener Capital Management Inc.5,000$236 03/31/2020
     IONS5,000$236 03/31/2020
Allianz Asset Management GmbH4,992$236 03/31/2020
     IONS4,992$236 03/31/2020
Stratos Wealth Partners LTD.4,752$225 03/31/2020
     IONS4,752$225 03/31/2020
Icon Wealth Partners LLC4,748$224 03/31/2020
     IONS4,748$224 03/31/2020
Regions Financial Corp4,567$216 03/31/2020
     IONS4,567$216 03/31/2020
Mirae Asset Global Investments Co. Ltd.4,521$214 03/31/2020
     IONS4,521$214 03/31/2020
Checchi Capital Advisers LLC4,340$205 03/31/2020
     IONS4,340$205 03/31/2020
Banque Cantonale Vaudoise4,350$205 03/31/2020
     IONS4,350$205 03/31/2020
Woodley Farra Manion Portfolio Management Inc.4,267$202 03/31/2020
     IONS4,267$202 03/31/2020
QS Investors LLC4,000$190 03/31/2020
     IONS4,000$190 03/31/2020
Steward Partners Investment Advisory LLC3,747$177 03/31/2020
     IONS3,747$177 03/31/2020
ETF Managers Group LLC3,395$164 03/31/2020
     IONS3,395$164 03/31/2020
American International Group Inc.2,910$138 03/31/2020
     IONS2,910$138 03/31/2020
Kistler Tiffany Companies LLC2,006$121 03/31/2020
     IONS2,006$121 03/31/2020
US Bancorp DE2,279$107 03/31/2020
     IONS2,279$107 03/31/2020
Panagora Asset Management Inc.2,200$104 03/31/2020
     IONS2,200$104 03/31/2020
Benjamin F. Edwards & Company Inc.2,089$99 03/31/2020
     IONS2,089$99 03/31/2020
Walled Lake Planning & Wealth Management LLC1,900$90 03/31/2020
     IONS1,900$90 03/31/2020
NEXT Financial Group Inc1,604$76 03/31/2020
     IONS1,604$76 03/31/2020
Schroder Investment Management Group1,392$71 03/31/2020
     IONS1,392$71 03/31/2020
Northwestern Mutual Wealth Management Co.1,518$71 03/31/2020
     IONS1,518$71 03/31/2020
Livforsakringsbolaget Skandia Omsesidigt1,500$71 03/31/2020
     IONS1,500$71 03/31/2020
S.A. Mason LLC1,510$71 03/31/2020
     IONS1,510$71 03/31/2020
Captrust Financial Advisors1,449$68 03/31/2020
     IONS1,449$68 03/31/2020
CSat Investment Advisory L.P.1,280$61 03/31/2020
     IONS1,280$61 03/31/2020
Allred Capital Management LLC1,271$60 03/31/2020
     IONS1,271$60 03/31/2020
First Manhattan Co.1,250$59 03/31/2020
     IONS1,250$59 03/31/2020
IFP Advisors Inc1,114$56 03/31/2020
     IONS1,114$56 03/31/2020
Acadian Asset Management LLC1,103$52 03/31/2020
     IONS1,103$52 03/31/2020
Institutional & Family Asset Management LLC1,032$49 03/31/2020
     IONS1,032$49 03/31/2020
Perkins Coie Trust Co1,000$47 03/31/2020
     IONS1,000$47 03/31/2020
Riggs Asset Managment Co. Inc.1,000$47 03/31/2020
     IONS1,000$47 03/31/2020
Bessemer Group Inc.900$43 03/31/2020
     IONS900$43 03/31/2020
Archer Investment Corp917$43 03/31/2020
     IONS917$43 03/31/2020
Harel Insurance Investments & Financial Services Ltd.892$42 03/31/2020
     IONS892$42 03/31/2020
Parallel Advisors LLC887$41 03/31/2020
     IONS887$41 03/31/2020
Glenmede Trust Co. NA646$38 03/31/2020
     IONS646$38 03/31/2020
Bradley Foster & Sargent Inc. CT804$38 03/31/2020
     IONS804$38 03/31/2020
Point72 Hong Kong Ltd791$37 03/31/2020
     IONS791$37 03/31/2020
FinTrust Capital Advisors LLC732$35 03/31/2020
     IONS732$35 03/31/2020
NWK Group Inc.750$35 03/31/2020
     IONS750$35 03/31/2020
Credit Agricole S A670$32 03/31/2020
     IONS670$32 03/31/2020
Signaturefd LLC653$31 03/31/2020
     IONS653$31 03/31/2020
Cornerstone Advisors Inc.663$31 03/31/2020
     IONS663$31 03/31/2020
Carroll Financial Associates Inc.641$30 03/31/2020
     IONS641$30 03/31/2020
Balentine LLC606$29 03/31/2020
     IONS606$29 03/31/2020
Laurel Wealth Advisors LLC600$28 03/31/2020
     IONS600$28 03/31/2020
Prestige Wealth Management Group LLC512$24 03/31/2020
     IONS512$24 03/31/2020
Quadrant Capital Group LLC490$23 03/31/2020
     IONS490$23 03/31/2020
Ellis Investment Partners LLC500$23 03/31/2020
     IONS500$23 03/31/2020
Jacobi Capital Management LLC427$20 03/31/2020
     IONS427$20 03/31/2020
Brown Brothers Harriman & Co.360$17 03/31/2020
     IONS360$17 03/31/2020
Pearl River Capital LLC343$16 03/31/2020
     IONS343$16 03/31/2020
Quantbot Technologies LP318$15 03/31/2020
     IONS318$15 03/31/2020
CWM LLC318$15 03/31/2020
     IONS318$15 03/31/2020
TFO TDC LLC325$15 03/31/2020
     IONS325$15 03/31/2020
Citizens Financial Group Inc RI298$14 03/31/2020
     IONS298$14 03/31/2020
Pathstone Family Office LLC282$14 03/31/2020
     IONS282$14 03/31/2020
Gables Capital Management Inc.275$13 03/31/2020
     IONS275$13 03/31/2020
Liberty Wealth Management LLC267$13 03/31/2020
     IONS267$13 03/31/2020
Synovus Financial Corp269$12 03/31/2020
     IONS269$12 03/31/2020
Firestone Capital Management250$12 03/31/2020
     IONS250$12 03/31/2020
Ameritas Investment Company LLC223$11 03/31/2020
     IONS223$11 03/31/2020
Cullen Frost Bankers Inc.212$10 03/31/2020
     IONS212$10 03/31/2020
Perennial Advisors LLC205$10 03/31/2020
     IONS205$10 03/31/2020
AllSquare Wealth Management LLC198$9 03/31/2020
     IONS198$9 03/31/2020
Reilly Financial Advisors LLC170$8 03/31/2020
     IONS170$8 03/31/2020
Winslow Evans & Crocker Inc.150$8 03/31/2020
     IONS150$8 03/31/2020
Pacifica Partners Inc.150$8 03/31/2020
     IONS150$8 03/31/2020
Shine Investment Advisory Services Inc.150$7 03/31/2020
     IONS150$7 03/31/2020
Bedel Financial Consulting Inc.138$7 03/31/2020
     IONS138$7 03/31/2020
Glassman Wealth Services124$6 03/31/2020
     IONS124$6 03/31/2020
Golden State Wealth Management LLC126$6 03/31/2020
     IONS126$6 03/31/2020
Center for Financial Planning Inc.135$6 03/31/2020
     IONS135$6 03/31/2020
Whittier Trust Co.100$5 03/31/2020
     IONS100$5 03/31/2020
Manchester Capital Management LLC101$5 03/31/2020
     IONS101$5 03/31/2020
Gradient Investments LLC109$5 03/31/2020
     IONS109$5 03/31/2020
Orion Portfolio Solutions LLC100$5 03/31/2020
     IONS100$5 03/31/2020
Blume Capital Management Inc.90$4 03/31/2020
     IONS90$4 03/31/2020
Global Wealth Management Investment Advisory Inc.80$4 03/31/2020
     IONS80$4 03/31/2020
CENTRAL TRUST Co60$3 03/31/2020
     IONS60$3 03/31/2020
LS Investment Advisors LLC57$3 03/31/2020
     IONS57$3 03/31/2020
TCI Wealth Advisors Inc.53$3 03/31/2020
     IONS53$3 03/31/2020
Wealthcare Advisory Partners LLC55$3 03/31/2020
     IONS55$3 03/31/2020
Trustcore Financial Services LLC58$3 03/31/2020
     IONS58$3 03/31/2020
Associated Banc Corp47$2 03/31/2020
     IONS47$2 03/31/2020
Meeder Asset Management Inc.42$2 03/31/2020
     IONS42$2 03/31/2020
Earnest Partners LLC49$2 03/31/2020
     IONS49$2 03/31/2020
Harbor Investment Advisory LLC38$2 03/31/2020
     IONS38$2 03/31/2020
Ellevest Inc.32$2 03/31/2020
     IONS32$2 03/31/2020
BDO Wealth Advisors LLC33$2 03/31/2020
     IONS33$2 03/31/2020
Versant Capital Management Inc46$2 03/31/2020
     IONS46$2 03/31/2020
Global Retirement Partners LLC41$2 03/31/2020
     IONS41$2 03/31/2020
Semmax Financial Advisors Inc.39$2 03/31/2020
     IONS39$2 03/31/2020
tru Independence LLC36$2 03/31/2020
     IONS36$2 03/31/2020
Enterprise Financial Services Corp11$1 03/31/2020
     IONS11$1 03/31/2020
AdvisorNet Financial Inc29$1 03/31/2020
     IONS29$1 03/31/2020
Cutler Group LP $1 03/31/2020
     Call1,000$1 03/31/2020
San Francisco Sentry Investment Group CA17$1 03/31/2020
     IONS17$1 03/31/2020
Allworth Financial LP24$1 03/31/2020
     IONS24$1 03/31/2020
Lowe Wealth Advisors LLC20$1 03/31/2020
     IONS20$1 03/31/2020
Cranbrook Wealth Management LLC25$1 03/31/2020
     IONS25$1 03/31/2020
Sowell Financial Services LLC30$1 03/31/2020
     IONS30$1 03/31/2020
Valeo Financial Advisors LLC15$1 03/31/2020
     IONS15$1 03/31/2020
Sound Income Strategies LLC20$1 03/31/2020
     IONS20$1 03/31/2020
Private Capital Group LLC31$1 03/31/2020
     IONS31$1 03/31/2020
Lindbrook Capital LLC20$1 03/31/2020
     IONS20$1 03/31/2020
Huntington National Bank1$0 03/31/2020
     IONS1$0 03/31/2020
Proequities Inc. $0 03/31/2020
     Put0$0 03/31/2020
     Call0$0 03/31/2020
     IONS0$0 03/31/2020
Gemmer Asset Management LLC3$0 03/31/2020
     IONS3$0 03/31/2020
Procyon Private Wealth Partners LLC3$0 03/31/2020
     IONS3$0 03/31/2020
Royal Fund Management LLC10$0 03/31/2020
     IONS10$0 03/31/2020
Wolverine Asset Management LLC $-85 03/31/2020
     Put1,800$85 03/31/2020
CSS LLC IL $-241 03/31/2020
     Put5,100$241 03/31/2020
Twin Tree Management LP $-412 03/31/2020
     Put9,300$440 03/31/2020
     IONS595$28 03/31/2020
Walleye Trading LLC $-714 03/31/2020
     Put65,100$3,078 03/31/2020
     IONS33,510$1,584 03/31/2020
     Call16,500$780 03/31/2020
BNP Paribas Arbitrage SA $-808 03/31/2020
     Put1,533$7,248 03/31/2020
     IONS72,350$3,421 03/31/2020
     BOND3,009,000$3,019 03/31/2020
Simplex Trading LLC $-1,909 03/31/2020
     Put725$3,427 03/31/2020
     Call261$1,234 03/31/2020
     IONS6,012$284 03/31/2020
PEAK6 Investments LLC16,647$-3,847 03/31/2020
     Put132,500$6,265 03/31/2020
     Call34,500$1,631 03/31/2020
     IONS16,647$787 03/31/2020

See Summary: Institutional Holders of IONS
See Details: Top 10 Hedge Funds Holding IONS
Also See: IONS Holdings Changes

Quotes delayed 20 minutes

Funds Holding IONS | www.HoldingsChannel.com

Copyright © 2013 - 2020, All Rights Reserved Nothing in Holdings Channel is intended to be investment advice, nor does it represent the opinion of, counsel from, or recommendations by BNK Invest Inc. or any of its affiliates, subsidiaries or partners. None of the information contained herein constitutes a recommendation that any particular security, portfolio, transaction, or investment strategy is suitable for any specific person. All viewers agree that under no circumstances will BNK Invest, Inc,. its subsidiaries, partners, officers, employees, affiliates, or agents be held liable for any loss or damage caused by your reliance on information obtained. By visiting, using or viewing this site, you agree to the following Full Disclaimer & Terms of Use and Privacy Policy. Video widget and videos powered by Market News Video. Quote data delayed at least 20 minutes; data powered by Ticker Technologies, and Mergent. Contact Holdings Channel; Meet Our Editorial Staff.